Environmental physiology and reproductive physiology of fish. Particularly, determination of effects of stress on fish performance in the wild and in the hatchery. Areas of interest include fish health, migration and development. Studies include synecology, endocrinology, immunology, molecular genetics as well as behavior and toxicology.
Ph.D. - 1972 - Physiology and Fish Biology, Colorado State Univ., Ft. Collins, CO
M.S. - 1969 - Fish Biology, Colorado State Univ., Ft. Collins,CO
B.S. - 1966 - Zoology, University of California, Berkeley, CA
Enjoys hiking, fly fishing, canoeing, skiing.